This study was carried out on nine ring-tailed lemurs and two red ruffed lemurs housed in a mixed exhibit at Furuvik Zoo, Sweden.

The lemurs’ enclosure at Furuvik Zoo comprises an outdoor area of 5,000 m², within which two indoor enclosures are located: the ring-tailed lemurs’ house (200 m³) and the red ruffed lemurs’ house (500 m³).

The enclosure’s outdoor area is covered in vegetation, including trees, bushes, shrubs, and grass.

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