This study took place from June to early October of 2008. A total of six locations in Sweden which hold European bisons were included in the study, five zoos and one breeding centre. Half of the study sites were of the naturalistic type and the other half of the barren type. The adult animals on these locations were observed continuously, using rotational focal sampling, for a total 96 hours per visit.  Each site was visited twice; the data collected during the first visit was used as a baseline and during the second visit extra browse was added and scattered to the degree possible, i.e. a lighter form of enrichment.  

Eight behaviours were recorded

• Feeding – active feeding on pasture or browse
• Feeding supl – feeding on hay or pellets
• Standing
• Standing rumi – standing and ruminating
• Walk
• Run
• Lying – lying down or wallowing
• Lying rumi – lying down ruminating