My gratitude goes to my supervisors; Lars Westerberg and Karl-Olof Bergman for their support and guidance throughout this work. I would like to also pass my appreciation to Ulrica Ronquist for all the field and lab work collaboration we executed together. Special thanks to Sven Lennartsson for his support in identifying beetle species, and Nicklas Jansson for practical and useful tips and essential input in planning of this study. I also thank Dennis Jonason for the thesis opportunity and support, and Anders Jorneskog for the information about the study area. A huge appreciation to the Swedish Institute for financing my studies. Finally, thanks to all fellow students who reviewed this work.